Brian M Wiegmann

Haplotype-resolved genome assemblies for the New World screwworm, Cochliomyia hominivorax (Diptera: Calliphoridae), using the trio binning approach

Hickner, P. V., Sim, S. B., Luecke, D., Bailey, E., Saelao, P., Yusseff-Vanegas, S. Z., … Olafson, P. U. (2026, February 28). G3 Genes Genomes Genetics, Vol. 2.

Phylogenomics of Anthomyiidae and a revised classification of Scathophagidae (Diptera: Calyptratae)

Bailey, E., Michelsen, V., Cassel, B., Szpila, K., Bayless, K., Grzywacz, A., … Wiegmann, B. (2026, January 30). DRYAD, Vol. 1.

Phylogenomics of Anthomyiidae and a revised classification of Scathophagidae (Diptera: Calyptratae)

Bailey, E. M., Michelsen, V., Cassel, B. K., Szpila, K., Bayless, K. M., Grzywacz, A., … Wiegmann, B. M. (2026, January 1). Systematic Entomology.

A Way Through the Trees: Molecular Phylogenies Consistently Recover Two Clades of Aedes Mosquitoes

Yeo, H., Chan, C., Morinaga, Wiegmann, B. M., & Soghigian, J. (2025, July 18).

A new phylogeny of Anastrepha (Diptera: Tephritidae) based on nuclear loci obtained by phylogenomic methods

Norrbom, A. L., Rodriguez, E. J., Steck, G. J., Cassel, B. K., Ruiz‐Arce, R., Muller, A., … Wiegmann, B. M. (2025, September 27). Systematic Entomology, Vol. 9.

Blow fly larvae socially integrate termite nests through morphological and chemical mimicry

Schär, S., Talavera, G., Dapporto, L., Bruschini, C., Dincă, V., Beza-Beza, C., … Vila, R. (2025, February 10). Current Biology, Vol. 35.

Phylogenomics and the evolution of larval feeding habits in the blow flies (Diptera: Calliphoridae)

Beza‐Beza, C. F., Soghigian, J., Bailey, E., Johnston, N. P., Cassel, B. K., Bayless, K. M., … Wiegmann, B. M. (2025, December 12). Systematic Entomology.

Population Phylogenomics and Genetic Structure of the Polyphagous Leafminer, Liriomyza trifolii (Burgess) (Diptera: Agromyzidae)

Xuan, J. L., Scheffer, S. J., Soghigian, J., Cassel, B., Lewis, M. L., Li, S. P., … Wiegmann, B. M. (2025, July 1). Evolutionary Applications, Vol. 18.

Chewing through challenges: Exploring the evolutionary pathways to wood‐feeding in insects

Beza‐Beza, C. F., Wiegmann, B. M., Ware, J. A., Petersen, M., Gunter, N., Cole, M. E., … Mikaelyan, A. (2024, March 27). BioEssays, Vol. 3.

Color Morphs in Anastrepha nigrotaenia (Enderlein), New Combination (Diptera: Tephritidae) and Resultant Synonymy

Norrbom, A. L., Moore, M. R., Paynter, Q., McGrath, Z., Probst, C. M., Korneyev, V. A., … Ruiz-Arce, R. (2024, August 22). Proceedings of the Entomological Society of Washington, Vol. 126, pp. 21–55.

Development of an agroinfiltration-based transient hairpin RNA expression system in pak choi leaves (Brassica rapa ssp. chinensis) for RNA interference against Liriomyza sativae

Li, S.-P., Chen, Z.-X., Gao, G., Bao, Y.-Q., Fang, W.-Y., Zhang, Y.-N., … Xuan, J.-L. (2024, August 19). Pesticide Biochemistry and Physiology, Vol. 204.

Anchored phylogenomics and revised classification of the Miltogramminae (Diptera: Sarcophagidae)

Johnston, N. P., Pape, T., Piwczyński, M., Wallman, J. F., Wiegmann, B. M., Cassel, B. K., … Szpila, K. (2023, October 10). Systematic Entomology, Vol. 10.

Chewing Through Challenges: Exploring the Evolutionary Pathways to Wood-Feeding in Insects

Beza-Beza, C. F., Wiegmann, B. M., Ware, J. A., Petersen, M., Gunter, N., Cole, M. E., … Mikaelyan, A. (2023, December 28).

New Host Plant and Distribution Records of Anastrepha Species (Diptera: Tephritidae) Primarily from the Western Amazon

Rodriguez, E. J., Norrbom, A. L., Steck, G. J., Moore, M. R., Sutton, B. D., Ruiz-Arce, R., … Branham, M. A. (2023, November 16). Proceedings of the Entomological Society of Washington, Vol. 125, pp. 89–164.

Phylogenomics reveals the history of host use in mosquitoes

Soghigian, J., Sither, C., Justi, S. A., Morinaga, Cassel, B. K., Vitek, C. J., … Wiegmann, B. M. (2023, October 6). Nature Communications, Vol. 14.

The phylogeny and divergence times of leaf-mining flies (Diptera: Agromyzidae) from anchored phylogenomics

Xuan, J.-L., Scheffer, S. J., Lewis, M., Cassel, B. K., Liu, W.-X., & Wiegmann, B. M. (2023, April 6). Molecular Phylogenetics and Evolution, Vol. 184.

A genome‐wide phylogeny and the diversification of genus Liriomyza (Diptera: Agromyzidae) inferred from anchored phylogenomics

Xuan, J. L., Scheffer, S. J., Lonsdale, O., Cassel, B. K., Lewis, M. L., Eiseman, C. S., … Wiegmann, B. M. (2022, September 15). Systematic Entomology, Vol. 9.

Phylogenetic resolution of the fly superfamily Ephydroidea–Molecular systematics of the enigmatic and diverse relatives of Drosophilidae

Winkler, I. S., Kirk-Spriggs, A. H., Bayless, K. M., Soghigian, J., Meier, R., Pape, T., … Wiegmann, B. M. (2022, October 5). PLoS ONE, Vol. 17.

Beyond Drosophila: resolving the rapid radiation of schizophoran flies with phylotranscriptomics

Bayless, K. M., Trautwein, M. D., Meusemann, K., Shin, S., Petersen, M., Donath, A., … Wiegmann, B. M. (2021, February 8). BMC Biology, Vol. 19.

Colonization with multidrug-resistant Enterobacteriaceae among infants: an observational study in southern Sri Lanka

Meredith, H. R., Kularatna, S., Nagaro, K., Nagahawatte, A., Bodinayake, C., Kurukulasooriya, R., … Tillekeratne, L. G. (2021, April 30). Antimicrobial Resistance and Infection Control, Vol. 4.

From e-voucher to genomic data: Preserving archive specimens as demonstrated with medically important mosquitoes (Diptera: Culicidae) and kissing bugs (Hemiptera: Reduviidae)

Justi, S. A., Soghigian, J., Pecor, D. B., Caicedo-Quiroga, L., Rutvisuttinunt, W., Li, T., … Linton, Y.-M. (2021, February 25). PLoS ONE, Vol. 16.

New species and host plants of Anastrepha (Diptera: Tephritidae) primarily from Suriname and Pará, Brazil

NORRBOM, A. L. L. E. N. L., Muller, A., GANGADIN, A. N. I. E. L. K. O. E. M. A. R., SUTTON, B. R. U. C. E. D., RODRIGUEZ, E. R. I. C. K. J., SAVARIS, M. A. R. C. O. A. N. D. R. E., … RUIZ-ARCE, R. A. U. L. (2021, September 26). Zootaxa, Vol. 9.

Towards a new classification of Muscidae (Diptera): a comparison of hypotheses based on multiple molecular phylogenetic approaches

Grzywacz, A., Trzeciak, P., Wiegmann, B. M., Cassel, B. K., Pape, T., Walczak, K., … Piwczyński, M. (2021, February 26). Systematic Entomology.

Are fleas highly modified Mecoptera? Phylogenomic resolution of Antliophora (Insecta: Holometabola)

Meusemann, K., Trautwein, M., Friedrich, F., Beutel, R. G., Wiegmann, B. M., Donath, A., … Yeates, D. K. (2020, November 20).

Phylogenomics reveals accelerated late Cretaceous diversification of bee flies (Diptera: Bombyliidae)

Li, X., Teasdale, L. C., Bayless, K. M., Ellis, A. G., Wiegmann, B. M., Lamas, C. J. E., … Yeates, D. K. (2020, November 20). Cladistics.

Anchored hybrid enrichment challenges the traditional classification of flesh flies (Diptera: Sarcophagidae)

Buenaventura, E., Szpila, K., Cassel, B. K., Wiegmann, B. M., & Pape, T. (2019, October 31). Systematic Entomology, Vol. 45, pp. 281–301.

First Record of Mansonia dyari From Saint Croix, United States Virgin Islands

Seger, K. R., Drummond, A., Delgado, D., Day, C. A., Sither, C. B., Soghigian, J., … Byrd, B. D. (2019, September 1). Journal of the American Mosquito Control Association, Vol. 35, pp. 214–216.

Phylogenomic analysis of Calyptratae: resolving the phylogenetic relationships within a major radiation of Diptera

Kutty, S. N., Meusemann, K., Bayless, K. M., Marinho, M. A. T., Pont, A. C., Zhou, X., … Pape, T. (2019, February 22). Cladistics, Vol. 35, pp. 605–622.

Structural and functional insights into the Diabrotica virgifera virgifera ATP-binding cassette transporter gene family

BMC Genomics, 20(1).

Anchored phylogenomics unravels the evolution of spider flies (Diptera, Acroceridae) and reveals discordance between nucleotides and amino acids

Gillung, J. P., Winterton, S. L., Bayless, K. M., Khouri, Z., Borowiec, M. L., Yeates, D., … Wiegmann, B. M. (2018, August 12). Molecular Phylogenetics and Evolution, Vol. 128, pp. 233–245.

Genomes of Diptera

Wiegmann, B. M., & Richards, S. (2018, February 1). Current Opinion in Insect Science, Vol. 25, pp. 116–124.

Taxon sampling to address an ancient rapid radiation: a supermatrix phylogeny of early brachyceran flies (Diptera)

SHIN, S. E. U. N. G. G. W. A. N., BAYLESS, K. E. I. T. H. M., WINTERTON, S. H. A. U. N. L., DIKOW, T. O. R. S. T. E. N., LESSARD, B. R. Y. A. N. D., YEATES, D. A. V. I. D. K., … TRAUTWEIN, M. I. C. H. E. L. L. E. D. (2017, November 11). Systematic Entomology, Vol. 43, pp. 277–289.

Advances using molecular data in insect systematics

Kjer, K., Borowiec, M. L., Frandsen, P. B., Ware, J., & Wiegmann, B. M. (2016, October 17). Current Opinion in Insect Science, Vol. 18, pp. 40–47.

Anchored enrichment dataset for true flies (order Diptera) reveals insights into the phylogeny of flower flies (family Syrphidae)

Young, A. D., Lemmon, A. R., Skevington, J. H., Mengual, X., Ståhls, G., Reemer, M., … Wiegmann, B. M. (2016, June 29). BMC Evolutionary Biology, Vol. 16.

Editorial overview: Insect phylogenetics: an expanding toolbox to resolve evolutionary questions

Current Opinion in Insect Science, 18, 93–95.

Mitochondrial Genome Sequences and Structures Aid in the Resolution of Piroplasmida phylogeny

Schreeg, M. E., Marr, H. S., Tarigo, J. L., Cohn, L. A., Bird, D. M., Scholl, E. H., … Birkenheuer, A. J. (2016, November 10). PLoS ONE, Vol. 11.

Molecular phylogeny of the horse flies: a framework for renewing tabanid taxonomy

MORITA, S. H. E. L. A. H. I., BAYLESS, K. E. I. T. H. M., YEATES, D. A. V. I. D. K., & WIEGMANN, B. R. I. A. N. M. (2015, September 26). Systematic Entomology, Vol. 41, pp. 56–72.

Power, resolution and bias: recent advances in insect phylogeny driven by the genomic revolution

Yeates, D. K., Meusemann, K., Trautwein, M., Wiegmann, B., & Zwick, A. (2015, November 14). Current Opinion in Insect Science, Vol. 13, pp. 16–23.

Response to Comment on "Phylogenomics resolves the timing and pattern of insect evolution"

Say goodbye to tribes in the new house fly classification: A new molecular phylogenetic analysis and an updated biogeographical narrative for the Muscidae (Diptera)

Haseyama, K. L. F., Wiegmann, B. M., Almeida, E. A. B., & Carvalho, C. J. B. (2015, April 11). Molecular Phylogenetics and Evolution, Vol. 89, pp. 1–12.

The phylogeny of stiletto flies ( D iptera: T herevidae)

WINTERTON, S. H. A. U. N. L., HARDY, N. A. T. E. B., GAIMARI, S. T. E. P. H. E. N. D., HAUSER, M. A. R. T. I. N., HILL, H. I. L. A. R. Y. N., HOLSTON, K. E. V. I. N. C., … WIEGMANN, B. R. I. A. N. M. (2015, October 6). Systematic Entomology, Vol. 41, pp. 144–161.

Where do the Neotropical Empidini lineages (Diptera: Empididae: Empidinae) fit in a worldwide context?

Watts, M., Winkler, I. S., Daugeron, C., Carvalho, C. J. B., Turner, S. P., & Wiegmann, B. M. (2015, November 29). Molecular Phylogenetics and Evolution, Vol. 95, pp. 67–78.

A revision of Afrotropical Quasimodo flies (Diptera: Schizophora; Curtonotidae). Part IV—the continental Afrotropical species of Curtonotum Macquart, with descriptions of thirteen new species and a combined phylogenetic analysis of the Curtonotidae

KIRK-SPRIGGS, A. S. H. L. E. Y. H., & WIEGMANN, B. R. I. A. N. M. (2013, July 8). Zootaxa, Vol. 3684, p. 1-.

Flies: The Natural History and Diversity of Diptera. By Stephen A. Marshall. Buffalo (New York): Firefly Books. $125.00. 616 p.; ill.; index. ISBN: 978-1-77085-100-9. 2012.

The Quarterly Review of Biology, 88(4), 359–359.

The evolution and biogeography of the austral horse fly tribe Scionini (Diptera: Tabanidae: Pangoniinae) inferred from multiple mitochondrial and nuclear genes

Lessard, B. D., Cameron, S. L., Bayless, K. M., Wiegmann, B. M., & Yeates, D. K. (2013, May 9). Molecular Phylogenetics and Evolution, Vol. 68, pp. 516–540.

A phylogenetic analysis of Sciomyzidae (Diptera) and some related genera

Tóthová, A., Rozkošný, R., Knutson, L., Kutty, S. N., Wiegmann, B. M., & Meier, R. (2012, November 2). Cladistics, Vol. 29, pp. 404–415.

The impact of theManual of Nearctic Dipteraon phylogenetic dipterology

Yeates, D. K., & Wiegmann, B. M. (2012, March 7). The Canadian Entomologist, Vol. 144, pp. 197–205.

The phylogenetic relationships among infraorders and superfamilies of Diptera based on morphological evidence

LAMBKIN, C. H. R. I. S. T. I. N. E. L., SINCLAIR, B. R. A. D. L. E. Y. J., PAPE, T. H. O. M. A. S., COURTNEY, G. R. E. G. W., SKEVINGTON, J. E. F. F. R. E. Y. H., MEIER, R. U. D. O. L. F., … WIEGMANN, B. R. I. A. N. M. (2012, September 25). Systematic Entomology, Vol. 38, pp. 164–179.

Advances in Insect Phylogeny at the Dawn of the Postgenomic Era

Trautwein, M. D., Wiegmann, B. M., Beutel, R., Kjer, K. M., & Yeates, D. K. (2011, March 3). Annual Review of Entomology, Vol. 57, p. 449-.

Episodic radiations in the fly tree of life

Proceedings of the National Academy of Sciences, 108(14), 5690–5695.

Episodic radiations in the fly tree of life

Proceedings of the National Academy of Sciences of the United States of America, 108(14), 5690–5695.

Rediscovery of the “terrible hairy fly”,Mormotomyia hirsutaAusten (Diptera: Mormotomyiidae), in Eastern Kenya, with Notes on Biology, Natural History, and Genetic Variation of the Ukasi Hill Population

Copeland, R. S., Kirk-Spriggs, A. H., Muteti, S., Booth, W., & Wiegmann, B. M. (2011, December 1). African Invertebrates, Vol. 52, pp. 363–390.

A multigene phylogeny of the fly superfamily Asiloidea (Insecta): Taxon sampling and additional genes reveal the sister-group to all higher flies (Cyclorrhapha)

Trautwein, M. D., Wiegmann, B. M., & Yeates, D. K. (2010, April 24). Molecular Phylogenetics and Evolution, Vol. 56, pp. 918–930.

Molecular phylogeny of the Calyptratae (Diptera: Cyclorrhapha) with an emphasis on the superfamily Oestroidea and the position of Mystacinobiidae and McAlpine's fly

KUTTY, S. U. J. A. T. H. A. N. A. R. A. Y. A. N. A. N., PAPE, T. H. O. M. A. S., WIEGMANN, B. R. I. A. N. M., & MEIER, R. U. D. O. L. F. (2010, August 5). Systematic Entomology, Vol. 35, pp. 614–635.

On wings of lace: phylogeny and Bayesian divergence time estimates of Neuropterida (Insecta) based on morphological and molecular data

WINTERTON, S. H. A. U. N. L., HARDY, N. A. T. E. B., & WIEGMANN, B. R. I. A. N. M. (2010, March 20). Systematic Entomology, Vol. 35, pp. 349–378.

Phylogenetic analysis of the druid flies (Diptera: Schizophora: Clusiidae) based on morphological and molecular data

Fu, J., Lonsdale, O., Wiegmann, B., & Marshall, S. (2010, January 1). Insect Systematics & Evolution, Vol. 41, pp. 231–274.

Phylogenetic synthesis of morphological and molecular data reveals new insights into the higher‐level classification of Tipuloidea (Diptera)

PETERSEN, M. A. T. T. H. E. W. J., BERTONE, M. A. T. T. H. E. W. A., WIEGMANN, B. R. I. A. N. M., & COURTNEY, G. R. E. G. O. R. Y. W. (2010, May 6). Systematic Entomology, Vol. 35, pp. 526–545.

Phylogenetic analysis and temporal diversification of mosquitoes (Diptera: Culicidae) based on nuclear genes and morphology

Reidenbach, K. R., Cook, S., Bertone, M. A., Harbach, R. E., Wiegmann, B. M., & Besansky, N. J. (2009, December 1). BMC Evolutionary Biology, Vol. 9.

Phylogenetic relationships of Ceratitis fruit flies inferred from nuclear CAD and tango/ARNT gene fragments: Testing monophyly of the subgenera Ceratitis (Ceratitis) and C. (Pterandrus)

Barr, N. B., & Wiegmann, B. M. (2009, July 15). Molecular Phylogenetics and Evolution, Vol. 53, pp. 412–424.

Single-copy nuclear genes resolve the phylogeny of the holometabolous insects

Wiegmann, B. M., Trautwein, M. D., Kim, J.-W., Cassel, B. K., Bertone, M. A., Winterton, S. L., & Yeates, D. K. (2009, June 24). BMC Biology, Vol. 7.

Supertrees and the Tree of Life: generating a metaphylogeny for a diverse invertebrate family (Insecta : Diptera : Therevidae) using constraint trees and the parsimony ratchet to overcome low taxon overlap

Lambkin, C. L., Trueman, J. W. H., Yeates, D. K., Holston, K. C., Webb, D. W., Hauser, M., … Wiegmann, B. M. (2009, June 4). Invertebrate Systematics, Vol. 23, pp. 171–191.

Molecular evolution of PISTILLATA-like genes in the dogwood genus Cornus (Cornaceae)

Zhang, W., Xiang, Q.-Y., Thomas, D. T., Wiegmann, B. M., Frohlich, M. W., & Soltis, D. E. (2008, January 2). Molecular Phylogenetics and Evolution, Vol. 47, pp. 175–195.

Phylogenetics and temporal diversification of the earliest true flies (Insecta: Diptera) based on multiple nuclear genes

BERTONE, M. A. T. T. H. E. W. A., COURTNEY, G. R. E. G. O. R. Y. W., & WIEGMANN, B. R. I. A. N. M. (2008, October 1). Systematic Entomology, Vol. 33, pp. 668–687.

The Muscoidea (Diptera: Calyptratae) are paraphyletic: Evidence from four mitochondrial and four nuclear genes

Kutty, S. N., Pape, T., Pont, A., Wiegmann, B. M., & Meier, R. (2008, August 30). Molecular Phylogenetics and Evolution, Vol. 49, pp. 639–652.

Molecular phylogenetics of the Muscidae (Diptera : Calyptratae): new ideas in a congruence context

Schuehli, G. S., Carvalho, C. J. B., & Wiegmann, B. M. (2007, June 26). Invertebrate Systematics, Vol. 21, pp. 263–278.

Monophyly and phylogenetic relationships of Thereva and therevine genus-groups (Insecta : Diptera : Therevidae) based on EF-1a, 28S rDNA and mitochondrial 16S rDNA sequences

Holston, K. C., Irwin, M. E., & Wiegmann, B. M. (2007, June 26). Invertebrate Systematics, Vol. 21, pp. 279–296.

Phylogenetic relationships within the leaf-mining flies (Diptera: Agromyzidae) inferred from sequence data from multiple genes

Scheffer, S. J., Winkler, I. S., & Wiegmann, B. M. (2007, January 1). Molecular Phylogenetics and Evolution, Vol. 42, pp. 756–775.

Phylogeny and systematics of Diptera: Two decades of progress and prospects

[Review of ]. Zootaxa, (1668), 565–590.

The phylogenetic relationships of flies in the superfamily Empidoidea (Insecta: Diptera)

Moulton, J. K., & Wiegmann, B. M. (2007, April 18). Molecular Phylogenetics and Evolution, Vol. 43, pp. 701–713.

The phylogeny and evolution of host choice in the Hippoboscoidea (Diptera) as reconstructed using four molecular markers

Petersen, F. T., Meier, R., Kutty, S. N., & Wiegmann, B. M. (2007, May 19). Molecular Phylogenetics and Evolution, Vol. 45, pp. 111–122.

Evocoidae (Diptera: Asiloidea), a new family name for Ocoidae, based on Evocoa , a replacement name for the Chilean genus Ocoa Yeates, Irwin, and Wiegmann 2003

Yeates, D. K., Irwin, M. E., & Wiegmann, B. M. (2006, February 10). Systematic Entomology, Vol. 31, p. 373.

Evolutionary patterns in the antR-Cor gene in the dwarf dogwood complex (Cornus, Cornaceae)

Genetica, 130(1), 19–34.

Phylogeny and Bayesian divergence time estimations of small-headed flies (Diptera: Acroceridae) using multiple molecular markers

Winterton, S. L., Wiegmann, B. M., & Schlinger, E. I. (2006, August 25). Molecular Phylogenetics and Evolution, Vol. 43, pp. 808–832.

Phylogenetic analysis of Themira (Sepsidae: Diptera): sensitivity analysis, alignment, and indel treatment in a multigene study

Laamanen, T. R., Meier, R., Miller, M. A., Hille, A., & Wiegmann, B. M. (2005, June 1). Cladistics, Vol. 21, pp. 258–271.

Heterogeneous evolution of the Myc-like Anthocyanin regulatory gene and its phylogenetic utility in Cornus L. (Cornaceae)

Fan, C., Purugganan, M. D., Thomas, D. T., Wiegmann, B. M., & Xiang, Q.-Y. (2004, October 5). Molecular Phylogenetics and Evolution, Vol. 33, pp. 580–594.

Phylogenetic analysis of the genus Thricops Rondani (Diptera: Muscidae) based on molecular and morphological characters

Savage, J., Wheeler, T. A., & Wiegmann, B. M. (2004, June 14). Systematic Entomology, Vol. 29, pp. 395–414.

Regarding the taxonomic status of Ophyra Robineau-Desvoidy (Diptera: Muscidae): A molecular approach

Zootaxa, (712), 12-.

Treehopper trees: phylogeny of Membracidae (Hemiptera: Cicadomorpha: Membracoidea) based on molecules and morphology

Cryan, J. R., Wiegmann, B. M., Deitz, L. L., Dietrich, C. H., & Whiting, M. F. (2004, October 1). Systematic Entomology, Vol. 29, pp. 441–454.

Evolution and phylogenetic utility of CAD (rudimentary) among Mesozoic-aged Eremoneuran Diptera (Insecta)

Moulton, J. K., & Wiegmann, B. M. (2003, October 9). Molecular Phylogenetics and Evolution, Vol. 31, pp. 363–378.

Ocoidae, a new family of asiloid flies (Diptera: Brachycera: Asiloidea), based on Ocoa chilensis gen. and sp.n. from Chile, South America

Yeates, D. K., Irwin, M. E., & Wiegmann, B. M. (2003, October 1). Systematic Entomology, Vol. 28, pp. 417–431.